How the Base Blockchain Ecosystem Works. Top 5 Applications
Base, a Layer 2 blockchain developed by Coinbase exchange, has become the largest L2 protocol by total value locked (TVL) in 2025, with over $4.35 billion in user deposits. A key differentiator is its lack of a native token, though its development team has confirmed plans to issue one. Base also led all L2s in cumulative revenue, generating over $80 million, and is second only to BNB Chain in monthly active users.
The article highlights five key applications in its ecosystem:
1. **Aerodrome**: The leading DEX on Base with $440 million in deposits, operating on a ve(3,3) model. It is the primary liquidity hub and plans to merge with Velodrome.
2. **Virtuals Protocol**: A platform for creating, tokenizing, and monetizing autonomous AI agents, forming an "agent economy" powered by its VIRTUAL token.
3. **Clanker**: An AI agent tool for instantly deploying memecoins via text prompts on Farcaster, having launched over 500,000 tokens and generating nearly $50 million in fees.
4. **Limitless**: The largest prediction market on Base, with over $550 million in volume, allowing users to bet on event outcomes.
5. **Zora**: A social network that transforms user content into tradable tokens on Base, with creators earning a share of trading fees. Its integration with the Base App has increased its prominence.
The ecosystem is noted for its high user activity and significant financial metrics, positioning Base as a major player in the L2 space.
RBK-crypto12/24 12:22